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i allmänna toxicitetsstudier omfattande ett flertal behandlingscykler utförda på råtta och apa var de viktigaste målorganen magtarmkanalen, vilket resulterade i kräkningar och/ eller diarré, hematopoetisk och lymfatisk vävnad som resulterade i cytopenier i perifert blod, atrofi i lymfvävnad, hematopoetisk hypocellularitet i benmärg, perifer neuropati (iakttagen hos apa, mus och hund) innefattande sensoriska axoner, samt lindriga njurförändringar.
in multi-cycle general toxicity studies conducted in the rat and monkey, the principal target organs included the gastrointestinal tract, resulting in vomiting and/ or diarrhoea, haematopoietic and lymphatic tissues resulting in peripheral blood cytopenias, lymphoid tissue atrophy and hematopoietic bone marrow hypocellularity: peripheral neuropathy (observed in monkeys, mice and dogs) involving sensory nerve axons; and mild changes in the kidneys.